Nippostrongylus smalesae sp. nov. (Nematoda: Heligmonellidae) Collected from Maxomys whiteheadi (Rodentia: Muridae) of Sumatra, Indonesia

Nippostrongylus smalesae sp. nov. (Nematoda: Heligmonellidae) is described from a murid Maxomys whiteheadi collectedin Riau, Sumatra, Indonesia. It is readily distinguished from all of the known congeners by having the ray 3 widelydiverged distally from the ray 4 in the left lobe of bursa copulatrix. This is the tenth named species of Nippostrongylus.The same species but with slight morphological difference was also found from M. whiteheadi and Niviventer cremoniventerin Kalimantan (Borneo Island), suggesting wide geographical distribution on the Sundashelf islands. Zootaxa, Vol. 4323, No. 4. Hal. 579-585